This was the response to a similar question from a Ram tech on another forum:
"The Mileage is stored in the Cluster and PCM but they get the mileage signal independently. The PCM mileage is used for emissions test calibrations and maintenance, AND can be adjusted to match cluster mileage by the dealer. Cluster mileage cannot be adjusted by the dealer and has to be sent to a
service center or a replacement cluster is sent with the mileage already burned in it. Yes the service center fires up the old one to check if its the same as the new one. Many lease customers are caught tampering with the mileage by checking the PCM mileage and warranty records as soon as the car is hooked up to the diagnostic machine the
VIN and mileage are sent to
chrysler."
Basically, if you replace a cluster with a used one, it will read whatever was on it from the previous vehicle. If you hook up a scan tool, it will show the actual mileage from the vehicle, stored in the PCM and possibly several other places. Its playing with fire unless properly documented. That's one way they catch people trying to tamper with mileage. The different modules which store the odometer info also changes periodically to make it harder for would-be hackers to cover their tracks.