If I'm reading the other forum correctly, it's the anti-drainback valve in the spin-on filter located inside the transmission. When you lookup transmission filter kits for the 545RFE, you'll see the standard flat filter we all are used to seeing from back in the day and a spin-on filter similar to the ones we use for engine oil. It's POSSIBLE that the check valve you're looking for is there. As far as I can see, there is no check valve in the transmission lines unless it is internal to the cooler.